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kson (2002) suggested that the turtle shell can function as a pH buffer. To endure through anoxic conditions, such as winter periods beneath ice or within anoxic mud at the bottom of ponds, turtles utilize two general physiological mechanisms: their shell releases carbonate buffers and uptakes lactic acid.[27] Respiration Respiration, for many amniotes, is achieved by the contraction and relaxation of specific muscle groups (i.e. intercostals, abdominal muscles, and/or a diaphragm) attached to an internal rib-cage that can expand or contract the body wall thus assisting airflow in and out of the lungs.[28] The ribs of Testudines, however, are fused with their carapace and external to their pelvic and pectoral girdles, a feature unique among turtles. This rigid shell is not capable of expansion. With their immobile rib-cage, Testudines have had to evolve special adaptations for respiration.[29][30] Turtle pulmonary ventilation occurs by using specific groups of abdominal muscles attached to their viscera and shell that pull the lungs ventrally during inspiration, where air is drawn in via a negative pressure gradient (Boyle's Law).[28] In expiration, the contraction of the transversus abdominis is the driving force by propelling the viscera into the lungs and expelling air under positive pressure.[29] Conversely, the relaxing and flattening of the oblique abdominis muscle pulls the transversus back down which, once again, draws air back into the lungs.[29] Important auxiliary muscles used for ventilatory processes are the pectoralis, which is used in conjunction with the transverse abdominis during inspiration, and the serratus, which moves with the abdominal oblique accompanying expiration. The lungs of Testudines are multi-chambered and attached their entire length down the carapace. The number of chambers can vary between taxa, though most commonly they have three lateral chambers, three medial chambers, and one terminal chamber.[31] As previously mentioned, the act of specific abdominal muscles pulling down the viscera (or pushing back up) is what allows for respiration in turtles. Specifically, it is the turtles large liver that pulls or pushes on the lungs.[29] Ventral to the lungs, in the coelomic cavity, the liver of turtles is attached directly to the right lung, and their stomach is directly attached to the left lung by the ventral mesopneumonium, which is attached to their liver by the ventral mesentery.[29] When the liver is pulled down, inspiration begins. Supporting the lungs is the post-pulmo











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<br /><span style=3D"color:#FFFFFF;font-size:5px;">
kson (2002) suggested that the turtle shell can function as a pH buffer. To=
 endure through anoxic conditions, such as winter periods beneath ice or wi=
thin anoxic mud at the bottom of ponds, turtles utilize two general physiol=
ogical mechanisms: their shell releases carbonate buffers and uptakes lacti=
c acid.[27]

Respiration
Respiration, for many amniotes, is achieved by the contraction and relaxati=
on of specific muscle groups (i.e. intercostals, abdominal muscles, and/or =
a diaphragm) attached to an internal rib-cage that can expand or contract t=
he body wall thus assisting airflow in and out of the lungs.[28] The ribs o=
f Testudines, however, are fused with their carapace and external to their =
pelvic and pectoral girdles, a feature unique among turtles. This rigid she=
ll is not capable of expansion. With their immobile rib-cage, Testudines ha=
ve had to evolve special adaptations for respiration.[29][30] Turtle pulmon=
ary ventilation occurs by using specific groups of abdominal muscles attach=
ed to their viscera and shell that pull the lungs ventrally during inspirat=
ion, where air is drawn in via a negative pressure gradient (Boyle's Law).[=
28] In expiration, the contraction of the transversus abdominis is the driv=
ing force by propelling the viscera into the lungs and expelling air under =
positive pressure.[29] Conversely, the relaxing and flattening of the obliq=
ue abdominis muscle pulls the transversus back down which, once again, draw=
s air back into the lungs.[29] Important auxiliary muscles used for ventila=
tory processes are the pectoralis, which is used in conjunction with the tr=
ansverse abdominis during inspiration, and the serratus, which moves with t=
he abdominal oblique accompanying expiration.

The lungs of Testudines are multi-chambered and attached their entire lengt=
h down the carapace. The number of chambers can vary between taxa, though m=
ost commonly they have three lateral chambers, three medial chambers, and o=
ne terminal chamber.[31] As previously mentioned, the act of specific abdom=
inal muscles pulling down the viscera (or pushing back up) is what allows f=
or respiration in turtles. Specifically, it is the turtles large liver that=
 pulls or pushes on the lungs.[29] Ventral to the lungs, in the coelomic ca=
vity, the liver of turtles is attached directly to the right lung, and thei=
r stomach is directly attached to the left lung by the ventral mesopneumoni=
um, which is attached to their liver by the ventral mesentery.[29] When the=
 liver is pulled down, inspiration begins. Supporting the lungs is the post=
-pulmo
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